diff --git a/modules/lora/native_adapter.py b/modules/lora/native_adapter.py
index 91b6e2d2b..72a330aa4 100644
--- a/modules/lora/native_adapter.py
+++ b/modules/lora/native_adapter.py
@@ -48,7 +48,11 @@ from modules.lora import lora_common as l
 # vendor-specific naming conventions. ``lora_transformer_`` is not a vendor
 # format but sdnext's own internal transformer namespace; files already saved
 # in it (e.g. OneTrainer) pass through verbatim, see :func:`resolve_group_targets`.
-KNOWN_PREFIXES_DEFAULT = ("diffusion_model.", "transformer.", "lora_unet_", "lora_transformer_")
+# ``lycoris_`` is the LyCORIS-standalone save format: the wrapped diffusers
+# module path with dots rendered as underscores. It is arch-independent by
+# construction (the wrapped layout equals the target layout), so it lives here
+# rather than in any arch's prefix list.
+KNOWN_PREFIXES_DEFAULT = ("diffusion_model.", "transformer.", "lora_unet_", "lora_transformer_", "lycoris_")
 
 
 # Sentinel ``prefix_used`` value emitted by :func:`parse_key` when a bare path
@@ -67,9 +71,9 @@ NETWORK_PREFIX_DEFAULT = "lora_transformer_"
 
 # Prefixes whose parsed ``base`` is already a network-key tail (``arch_prefix +
 # base.replace(".", "_")`` matches the stamped module name), so the loader binds
-# them directly with no per-arch rewrite. Arch-local already-resolved prefixes
-# (e.g. flux2's ``lycoris_``) stay in that arch's ``resolve_targets``.
-PASSTHROUGH_PREFIXES_DEFAULT = ("transformer.", BARE_DIFFUSERS_PREFIX_USED, "lora_transformer_")
+# them directly with no per-arch rewrite. ``lycoris_`` bases are already
+# underscored, so the loader's ``.replace(".", "_")`` is a no-op on them.
+PASSTHROUGH_PREFIXES_DEFAULT = ("transformer.", BARE_DIFFUSERS_PREFIX_USED, "lora_transformer_", "lycoris_")
 
 
 def _resolve_prefix(network_prefix, prefix_used):
diff --git a/pipelines/anima/anima_lora.py b/pipelines/anima/anima_lora.py
index cc9c01dbf..cea98faa8 100644
--- a/pipelines/anima/anima_lora.py
+++ b/pipelines/anima/anima_lora.py
@@ -51,6 +51,10 @@ ANIMA_PREFIXES = (
     "diffusion_model.",
     "lora_te_",
     "lora_unet_",
+    # LyCORIS-standalone save format; resolved by the universal passthrough in
+    # native_adapter.resolve_group_targets and routed to lora_transformer_ by
+    # network_prefix_for's default arm.
+    "lycoris_",
 )
 
 
diff --git a/pipelines/flux/flux2_lora.py b/pipelines/flux/flux2_lora.py
index 9754e71b3..681c65087 100644
--- a/pipelines/flux/flux2_lora.py
+++ b/pipelines/flux/flux2_lora.py
@@ -56,7 +56,7 @@ from modules.lora.native_adapter import ChunkSpec
 
 # === Arch-specific prefix configuration ===
 
-KNOWN_PREFIXES = native_adapter.KNOWN_PREFIXES_DEFAULT + ("lycoris_",)
+KNOWN_PREFIXES = native_adapter.KNOWN_PREFIXES_DEFAULT
 
 BARE_FLUX_PREFIXES = (
     "single_blocks.", "double_blocks.", "img_in.", "txt_in.",
@@ -183,25 +183,16 @@ def resolve_targets(prefix_used, base):
     """Return ``[(diffusers_path, ChunkSpec | None), ...]`` for a parsed group key.
 
     For ``lora_unet_`` prefix, applies ``KOHYA_SUFFIX_MAP`` then ``F2_*_MAP``.
-    For BFL / bare-BFL, applies ``F2_*_MAP`` directly. ``lycoris_`` is an
-    already-underscored diffusers path, returned verbatim. Unrecognized
-    prefixes return an empty list.
+    For BFL / bare-BFL, applies ``F2_*_MAP`` directly. Unrecognized prefixes
+    return an empty list.
 
-    Universal passthrough prefixes are handled upstream by
-    :func:`native_adapter.resolve_group_targets`.
+    Universal passthrough prefixes (including ``lycoris_``) are handled
+    upstream by :func:`native_adapter.resolve_group_targets`.
     """
     if prefix_used == "lora_unet_":
         return _kohya_to_diffusers_targets(base)
     if prefix_used in (None, "diffusion_model."):
         return _bfl_to_diffusers_targets(base)
-    if prefix_used == "lycoris_":
-        # base is an already-underscored diffusers path (e.g.
-        # 'transformer_blocks_0_attn_add_k_proj'). The caller's network_key
-        # construction does base.replace('.', '_'); for already-underscored
-        # paths that's a no-op, so the network_key matches the entry stamped
-        # by lora_convert.assign_network_names_to_compvis_modules
-        # (e.g. 'lora_transformer_transformer_blocks_0_attn_add_k_proj').
-        return [(base, None)]
     return []
 
 
diff --git a/test/test-flux2-native-adapters.py b/test/test-flux2-native-adapters.py
index a595f529d..162774d42 100644
--- a/test/test-flux2-native-adapters.py
+++ b/test/test-flux2-native-adapters.py
@@ -718,8 +718,9 @@ def test_parse_key_lycoris_prefix():
         got = F.parse_key(key, suffixes)
         assert got == expected, f'parse_key({key!r}) = {got}, expected {expected}'
 
-    # resolve_targets: the underscored path is returned verbatim (no chunk).
-    targets = F.resolve_targets('lycoris_', 'transformer_blocks_0_attn_add_k_proj')
+    # Resolution: lycoris_ is a universal passthrough handled upstream of the
+    # arch resolver, so the underscored path is returned verbatim (no chunk).
+    targets = F.native_adapter.resolve_group_targets(F.resolve_targets, 'lycoris_', 'transformer_blocks_0_attn_add_k_proj')
     assert targets == [('transformer_blocks_0_attn_add_k_proj', None)], f'targets={targets}'
     return True
 
diff --git a/test/test-zimage-native-adapters.py b/test/test-zimage-native-adapters.py
index 6f958161b..9e7a29a0e 100644
--- a/test/test-zimage-native-adapters.py
+++ b/test/test-zimage-native-adapters.py
@@ -313,6 +313,24 @@ def sd_lokr_legacy_fused_qkv():
     }
 
 
+def sd_lokr_lycoris_style():
+    """LyCORIS-standalone LoKR (``lycoris_`` + underscored diffusers path).
+
+    The prefix is resolved by the universal passthrough in native_adapter, not
+    by anything zimage-specific. ``to_out.0`` exercises the verbatim
+    round-trip where a naive underscore-to-dot expansion would corrupt the
+    ModuleList index.
+    """
+    return {
+        'lycoris_layers_0_attention_to_q.lokr_w1': torch.randn(LOKR_W1_DIM, LOKR_W1_DIM),
+        'lycoris_layers_0_attention_to_q.lokr_w2': torch.randn(HIDDEN // LOKR_W1_DIM, HIDDEN // LOKR_W1_DIM),
+        'lycoris_layers_0_attention_to_q.alpha': torch.tensor(float(LOKR_W1_DIM)),
+        'lycoris_layers_1_attention_to_out_0.lokr_w1': torch.randn(LOKR_W1_DIM, LOKR_W1_DIM),
+        'lycoris_layers_1_attention_to_out_0.lokr_w2': torch.randn(HIDDEN // LOKR_W1_DIM, HIDDEN // LOKR_W1_DIM),
+        'lycoris_layers_1_attention_to_out_0.alpha': torch.tensor(float(LOKR_W1_DIM)),
+    }
+
+
 def sd_loha_bfl_proj():
     """LoHA on attention.to_out.0 (non-fused; LoHA on fused qkv is skipped by the loader)."""
     return {
@@ -599,6 +617,20 @@ def test_lokr_legacy_fused_qkv_chunked():
     return True
 
 
+def test_lokr_lycoris_prefix_passthrough():
+    """lycoris_ keys load via the universal passthrough (hoisted, not zimage-specific)."""
+    net = _load_via(Z.try_load_lokr, sd_lokr_lycoris_style())
+    assert net is not None and len(net.modules) == 2, f'got {net.modules if net else None}'
+    expected = {
+        'lora_transformer_layers_0_attention_to_q',
+        'lora_transformer_layers_1_attention_to_out_0',
+    }
+    assert set(net.modules) == expected, f'got {set(net.modules)}'
+    for mod in net.modules.values():
+        assert isinstance(mod, network_lokr.NetworkModuleLokr)
+    return True
+
+
 def test_loha_bfl_proj():
     """BFL LoHA on a non-fused proj target binds via NetworkModuleHada."""
     net = _load_via(Z.try_load_loha, sd_loha_bfl_proj())
@@ -729,6 +761,7 @@ def run_tests():
         test_lora_dora_threading,
         test_lokr_bfl_adaln,
         test_lokr_legacy_fused_qkv_chunked,
+        test_lokr_lycoris_prefix_passthrough,
         test_loha_bfl_proj,
         test_loha_legacy_fused_qkv_chunked,
         test_oft_lycoris_no_npe,